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He was elected to Parliament at the 1923 elections for the constituency of Rodope, initially for the Democratic Union, and later founded the National Republican Party (Εθνικό Δημοκρατικό Κόμμα), renamed in 1928 National Radical Party (Εθνικό Ριζοσπαστικό Κόμμα). He was war minister from March to June 1924. On 24 August 1926, he overthrew the dictatorship of Theodoros Pangalos in a bloodless coup and formed a government, proclaiming elections for November. Notably, his party did not participate in these. In the elections of August 1928, voters elected nine of his party's candidates as MPs, and he was elected in Kavala.
During this time, Kondylis began moving rightward. In 1932 he became war minister again in return for his support of the Populist government, a post he retained after the Populists were reelected in 1933. From this post he was instrumental in crushing the March 1935 Venizelist revolt. In the period immediately following the revolt, Kondylis became the real power in the country. He sacked numerous pro-republican soldiers and civil servants, and condemned Venizelos to death in absentia.
By now, Kondylis was one of the strongest proponents of restoring the monarchy. However, he opposed Prime Minister Panagis Tsaldaris' call for a referendum. On 10 October 1935, Kondylis and several other officers called on Tsaldaris and forced him to resign. Kondylis forced President Alexandros Zaimis to name him the new premier. Later that day, Kondylis forced Zaimis to resign, declared himself Regent, abolished the Republic and staged a plebiscite on 3 November for the return of the monarchy.
The official tally showed that 98 percent of the voters supported the return of George II—an suspiciously high total that was likely obtained through fraud. Indeed, the vote took place under less-than-secret conditions. Voters were given the choice of dropping a blue piece of paper in the ballot box if they supported the monarchy, and a red one if they supported the republic. Those who supported the republic risked being beaten up. Under those circumstances, it took a brave Greek to vote "no". By this time, Kondylis had turned so far to the right that he now openly sympathized with fascism. He hoped to echo Benito Mussolini's example in Italy, in which Victor Emmanuel III had been reduced to a puppet.[2]
George returned to Greece on 25 November, and retained Kondylis as prime minister. Kondylis soon quarreled with the king, who was not content to be a mere puppet, and resigned five days later. In the January 1936 elections, he cooperated with Ioannis Rallis and managed to have fifteen MPs elected. Soon after, however, he died of a heart attack on 1 February 1936, in Athens. His nephew, Georgios Kondylis Jr., became a general in the Hellenic army and later fought against the Axis during the German invasion of Greece.